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The last of the Divine Generals have fallen and all that stands before the Great Gurren Brigade and victory, is the Imperial City Teppelin. However right before their eyes Teppelin collapses and reveals its true form, as a Gunman of gigantic proportions. Nia request to go along with Simon, in the Gurren-Lagann, to confront her father Lord Genome. First they must penetrate the impregnable defense.